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Lipidomic analysis revealed that GSH depletion significantly decreased levels of neutral lipids, including diacylglycerols (DAGs), triacylglycerols (TAGs), and cholesterol esters (CEs), while GSH depletion increased levels of sphingolipids and phospholipids (Figure 4, B and C, and Supplemental Figure 3, A–C), many of which have been shown to improve hepatic steatosis (26)

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The pooled phage output surviving PACE on PC_SS1 was next subjected to phage-assisted non-continuous evolution (PANCE) 52,53,54 on PC_SS2, which contained six additional mutations toward procaspase-1 (Fig